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Anti-tank vehicles made their first major appearance in World War II. As combatants developed effective armoured vehicles and tactics to use them, there were a number of experiments at producing vehicles specifically designed to stop enemy tanks.
These tank desroyers fell broadly into two categories. Some were designed to be faster and cheaper than a medium tank while still able to destroy heavy armour at long range. The second design option was to create a heavily-armoured vehicle that was more effective in armoured combat than the enemy tanks.
In German and Soviet designs the traditional turret was removed from an existing tank design, and a larger gun was mounted with a limited traverse in the hull. The weight and space savings of removing the turret allowed a smaller chassis to carry a larger gun. This was used by both forces to keep older designs competitive in the rapidly up-armoring of all AFV's that took place during the war.
For instance, the German Panzer I was obsolete before the war even started, with thin armor and only machine guns for armament. Yet they were forced into battle during the invasion of Poland, where they were found to be deathtraps. Before the invasion of France, 202 were rebuilt with a Czech 47 mm gun, becoming the Panzerjäger I. Soon the same sort of thing happened to the Panzer II during Operation Barbarossa: crews took to using captured Soviet 76.2mm anti-tank guns on them in various makeshift mountings in order to allow them stand off from their opposition. In this form they were later modified at the factory, producing the Marder II. Whereas the Panzerjäger concept used the chassis of obsolescent tanks, dedicated Jagdpanzer versions of tanks were developed later in the war. The Jagdpanther version of the Panther tank is considered the best of the Jagdpanzers.
The resulting US designs retained the turret, but left it open on top for more working room with the larger gun. The larger guns required a weight to be added to the rear of the turret, which can be seen on designs like the M10. The open top made them particularly vunerable to even handguns, and the very idea of independent anti-tank groups was found unworkable. By 1944 the "basic" Sherman was being upgunned in the field to be a Sherman Firefly, and by the end of that year heavier guns were being mounted on production Shermans.