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Ronia, its main character, is a girl growing up in a castle in the wood in a band of robbers; as the chief's only child, she is expected to become the leader of the band someday. Their castle, Matt's Fort, was split in two parts by a lightning bolt on the day of her birth; soon afterwards, a different robber clan settled one section, resulting in perpetual armed strife. Ronia feels rather out of place in this harsh environment; nearly her only friend is the old man Noddle-Pete, and for a while she hates her father Matt. One day, Ronia crawls through a tunnel to the other side, and meets Birk Borkason, the only son of the enemy chieftain, Borka. Although Ronia knows he belongs to the hated enemy clan, and initially hates him for it, she ends up becoming fast friends with him; ultimately, they both forsake their feuding families and call each other sister and brother. In the end, their families repent of their thieving and feuding, and everyone is reunited.
The story contains several made-up creatures, such as goblin folk, gray dwarfs and the cruel but beautiful wild harpies.
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