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Originally, planes were used for reconnisance and observation until somebody brought a rifle and took a potshot at another plane.
Guns have been used in planes until quite recently, when they were replaced by guided missiles such as the Sidewinder.
Even the US F/A-22 Raptor, due to enter service in 2005, has a gatling-style gun, but it only carries about 480 rounds of ammunition.
Military stubs Mil-stub